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Buildings are cards that, like Spells and Summons can only be played once the game has reached a certain time, depending on their Stage and paying with Mana 
.
The main difference to Summons is the fact that Buildings don't move.
Buildings can only be placed within the green spawning areas, in range of your Nexus or one of your Lanetowers.
Many Buildings have the ability Lifetime: Building, limiting their time on the field, even when you place them out of reach for enemies.